About the Authors

Yangyang Qiao

Contributed equally to this work with: Yangyang Qiao, Zhitao Wang

Roles Data curation, Formal analysis, Investigation, Validation, Visualization

Affiliation Division of Avian Infectious Diseases, State Key Laboratory of Veterinary Biotechnology, Harbin Veterinary Research Institute, the Chinese Academy of Agricultural Sciences, Harbin, the People’s Republic of China

Zhitao Wang

Contributed equally to this work with: Yangyang Qiao, Zhitao Wang

Roles Data curation, Formal analysis, Investigation, Validation, Visualization, Writing – original draft, Writing – review & editing

Affiliations Division of Avian Infectious Diseases, State Key Laboratory of Veterinary Biotechnology, Harbin Veterinary Research Institute, the Chinese Academy of Agricultural Sciences, Harbin, the People’s Republic of China, College of Veterinary Medicine, Northeast Agricultural University, Harbin, the People’s Republic of China

Zongxi Han

Roles Validation

Affiliation Division of Avian Infectious Diseases, State Key Laboratory of Veterinary Biotechnology, Harbin Veterinary Research Institute, the Chinese Academy of Agricultural Sciences, Harbin, the People’s Republic of China

Yuhao Shao

Roles Investigation, Validation

Affiliation Division of Avian Infectious Diseases, State Key Laboratory of Veterinary Biotechnology, Harbin Veterinary Research Institute, the Chinese Academy of Agricultural Sciences, Harbin, the People’s Republic of China

Yong Ma

Roles Investigation, Validation

Affiliation Division of Avian Infectious Diseases, State Key Laboratory of Veterinary Biotechnology, Harbin Veterinary Research Institute, the Chinese Academy of Agricultural Sciences, Harbin, the People’s Republic of China

Yumeng Liang

Roles Investigation, Validation

Affiliation Division of Avian Infectious Diseases, State Key Laboratory of Veterinary Biotechnology, Harbin Veterinary Research Institute, the Chinese Academy of Agricultural Sciences, Harbin, the People’s Republic of China

Zhijie Chen

Roles Investigation, Validation

Affiliations Division of Avian Infectious Diseases, State Key Laboratory of Veterinary Biotechnology, Harbin Veterinary Research Institute, the Chinese Academy of Agricultural Sciences, Harbin, the People’s Republic of China, College of Veterinary Medicine, Northeast Agricultural University, Harbin, the People’s Republic of China

Hanguang Wu

Roles Investigation, Validation

Affiliation Division of Avian Infectious Diseases, State Key Laboratory of Veterinary Biotechnology, Harbin Veterinary Research Institute, the Chinese Academy of Agricultural Sciences, Harbin, the People’s Republic of China

Lu Cui

Roles Investigation, Validation

Affiliation Division of Avian Infectious Diseases, State Key Laboratory of Veterinary Biotechnology, Harbin Veterinary Research Institute, the Chinese Academy of Agricultural Sciences, Harbin, the People’s Republic of China

Yanhui Zhang

Roles Investigation, Validation

Affiliation Division of Avian Infectious Diseases, State Key Laboratory of Veterinary Biotechnology, Harbin Veterinary Research Institute, the Chinese Academy of Agricultural Sciences, Harbin, the People’s Republic of China

Shengwang Liu

Roles Conceptualization, Data curation, Formal analysis, Funding acquisition, Project administration, Supervision, Writing – original draft, Writing – review & editing

liushengwang@caas.cn (SL); lihai@caas.cn (HL)

Affiliation Division of Avian Infectious Diseases, State Key Laboratory of Veterinary Biotechnology, Harbin Veterinary Research Institute, the Chinese Academy of Agricultural Sciences, Harbin, the People’s Republic of China

Hai Li

Roles Conceptualization, Data curation, Formal analysis, Funding acquisition, Project administration, Supervision, Writing – original draft, Writing – review & editing

liushengwang@caas.cn (SL); lihai@caas.cn (HL)

Affiliation Division of Avian Infectious Diseases, State Key Laboratory of Veterinary Biotechnology, Harbin Veterinary Research Institute, the Chinese Academy of Agricultural Sciences, Harbin, the People’s Republic of China

Competing Interests

The authors have declared that no competing interests exist.