About the Authors

Yoshihisa Ueno

Contributed equally to this work with: Yoshihisa Ueno, Riichiro Yoshida

Current address: Department of Agriculture, Ryukoku University, Seta Ohe-cho, Otsu-shi, Shiga, Japan

Affiliation Disease Resistant Crops Research Unit, National Institute of Agrobiological Sciences, Tsukuba, Ibaraki, Japan

Riichiro Yoshida

Contributed equally to this work with: Yoshihisa Ueno, Riichiro Yoshida

Current address: Faculty of Agriculture, Kagoshima University, Kagoshima, Japan

Affiliation Disease Resistant Crops Research Unit, National Institute of Agrobiological Sciences, Tsukuba, Ibaraki, Japan

Mitsuko Kishi-Kaboshi

Current address: National Institute of Floricultural Science, Tsukuba, Ibaraki, Japan

Affiliation Disease Resistant Crops Research Unit, National Institute of Agrobiological Sciences, Tsukuba, Ibaraki, Japan

Akane Matsushita

Current address: Du Pont Kabushiki Kaisha, Chiyoda-ku, Tokyo, Japan

Affiliation Disease Resistant Crops Research Unit, National Institute of Agrobiological Sciences, Tsukuba, Ibaraki, Japan

Chang-Jie Jiang

Affiliation Disease Resistant Crops Research Unit, National Institute of Agrobiological Sciences, Tsukuba, Ibaraki, Japan

Shingo Goto

Current address: Okitsu Citrus Research Station, National Institute of Fruit Tree Science, Shimizu, Shizuoka, Japan

Affiliation Disease Resistant Crops Research Unit, National Institute of Agrobiological Sciences, Tsukuba, Ibaraki, Japan

Akira Takahashi

Affiliation Disease Resistant Crops Research Unit, National Institute of Agrobiological Sciences, Tsukuba, Ibaraki, Japan

Hirohiko Hirochika

Affiliation Disease Resistant Crops Research Unit, National Institute of Agrobiological Sciences, Tsukuba, Ibaraki, Japan

Hiroshi Takatsuji

takatsuh@affrc.go.jp

Affiliation Disease Resistant Crops Research Unit, National Institute of Agrobiological Sciences, Tsukuba, Ibaraki, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: YU RY AM CJJ SG HT. Performed the experiments: YU RY. Analyzed the data: YU RY HT. Contributed reagents/materials/analysis tools: MKK AT HH. Wrote the paper: YU HT.