About the Authors

Lee Kozakiewicz

Contributed equally to this work with: Lee Kozakiewicz, Yong Chen

Affiliation Department of Microbiology and Immunology, Albert Einstein College of Medicine, the Bronx, New York, New York, United States of America

Yong Chen

Contributed equally to this work with: Lee Kozakiewicz, Yong Chen

Affiliation Department of Microbiology and Immunology, Albert Einstein College of Medicine, the Bronx, New York, New York, United States of America

Jiayong Xu

Affiliation Department of Microbiology and Immunology, Albert Einstein College of Medicine, the Bronx, New York, New York, United States of America

Yanhua Wang

Affiliation Department of Pathology, Albert Einstein College of Medicine, the Bronx, New York, New York, United States of America

Kyri Dunussi-Joannopoulos

Affiliation Immunology and Autoimmunity, Biotherapeutic Research, Pfizer, Cambridge, Massachusetts, United States of America

Qinglin Ou

Affiliation Department of Immunology and Antibody Engineering, Genentech, San Francisco, California, United States of America

Joanne L. Flynn

Affiliation Department of Microbiology and Molecular Genetics, University of Pittsburgh School of Medicine, Pittsburgh, Pennsylvania, United States of America

Steven A. Porcelli

Affiliations Department of Microbiology and Immunology, Albert Einstein College of Medicine, the Bronx, New York, New York, United States of America, Department of Medicine, Albert Einstein College of Medicine, Bronx, New York, United States of America

William R. Jacobs Jr

Affiliations Department of Microbiology and Immunology, Albert Einstein College of Medicine, the Bronx, New York, New York, United States of America, Howard Hughes Medical Institute, Albert Einstein College of Medicine, the Bronx, New York, New York, United States of America

John Chan

john.chan@einstein.yu.edu

Affiliations Department of Microbiology and Immunology, Albert Einstein College of Medicine, the Bronx, New York, New York, United States of America, Department of Medicine, Albert Einstein College of Medicine, Bronx, New York, United States of America

Competing Interests

Kyri Dunussi-Joannopoulos was affiliated as a researcher with Wyeth/Pfizer. Qinglin Ou is currently a researcher at Genentech. This does not alter our adherence to all PLoS Pathogens policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: LK YC KDJ QO JLF SAP WRJ JC. Performed the experiments: LK YC JX YW. Analyzed the data: LK YC YW JLF SAP WRJ JC. Contributed reagents/materials/analysis tools: KDJ QO. Wrote the paper: LK YC JLF JC.