About the Authors

Alex T. Nielsen

Affiliation Department of Microbiology and Immunology, Stanford University School of Medicine, Stanford, California, United States of America

Nadia A. Dolganov

Affiliation Department of Microbiology and Immunology, Stanford University School of Medicine, Stanford, California, United States of America

Thomas Rasmussen

Affiliation Technical University of Denmark, Department of Systems Biology, Kgs. Lyngby, Denmark

Glen Otto

Affiliation Department of Comparative Medicine, Stanford University School of Medicine, Stanford, California, United States of America

Michael C. Miller

Affiliation Department of Microbiology and Immunology, Stanford University School of Medicine, Stanford, California, United States of America

Stephen A. Felt

Affiliation Department of Comparative Medicine, Stanford University School of Medicine, Stanford, California, United States of America

Stéphanie Torreilles

Affiliation Department of Comparative Medicine, Stanford University School of Medicine, Stanford, California, United States of America

Gary K. Schoolnik

Gary.Schoolnik@stanford.edu

Affiliation Department of Microbiology and Immunology, Stanford University School of Medicine, Stanford, California, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: ATN NAD GKS. Performed the experiments: ATN NAD TR GO SAF ST. Analyzed the data: ATN NAD. Contributed reagents/materials/analysis tools: ATN NAD MCM. Wrote the paper: ATN GKS. Involved in performing all rabbit experiments and transcriptional analysis, and the first to identify the localized and significant expression of virulence genes in the mucus during infection: NAD.