About the Authors

Anna L. Tyler

Affiliation The Jackson Laboratory, Bar Harbor, Maine, United States of America

Leah Rae Donahue

Affiliation The Jackson Laboratory, Bar Harbor, Maine, United States of America

Gary A. Churchill

Affiliation The Jackson Laboratory, Bar Harbor, Maine, United States of America

Gregory W. Carter

Greg.Carter@jax.org

Affiliation The Jackson Laboratory, Bar Harbor, Maine, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: ALT LRD GAC GWC. Performed the experiments: LRD. Analyzed the data: ALT GAC GWC. Contributed reagents/materials/analysis tools: LRD GAC GWC. Wrote the paper: ALT GAC GWC.