About the Authors

Tatsuya Komura-Kawa

Affiliation Graduate School of Life and Environmental Sciences, University of Tsukuba, Tsukuba, Ibaraki, Japan

Keiko Hirota

Affiliations Graduate School of Life and Environmental Sciences, University of Tsukuba, Tsukuba, Ibaraki, Japan, Life Science Center, Tsukuba Advanced Research Alliance, University of Tsukuba, Tsukuba, Ibaraki, Japan

Yuko Shimada-Niwa

Affiliation Life Science Center, Tsukuba Advanced Research Alliance, University of Tsukuba, Tsukuba, Ibaraki, Japan

Rieko Yamauchi

Affiliations Graduate School of Life and Environmental Sciences, University of Tsukuba, Tsukuba, Ibaraki, Japan, Life Science Center, Tsukuba Advanced Research Alliance, University of Tsukuba, Tsukuba, Ibaraki, Japan

MaryJane Shimell

Affiliation Department of Genetics, Cell Biology and Development, University of Minnesota, Minneapolis, Minnesota, United States of America

Tetsuro Shinoda

Affiliation National Institute of Agrobiological Sciences, Tsukuba, Ibaraki, Japan

Akiyoshi Fukamizu

Affiliations Graduate School of Life and Environmental Sciences, University of Tsukuba, Tsukuba, Ibaraki, Japan, Life Science Center, Tsukuba Advanced Research Alliance, University of Tsukuba, Tsukuba, Ibaraki, Japan

Michael B. O’Connor

Affiliation Department of Genetics, Cell Biology and Development, University of Minnesota, Minneapolis, Minnesota, United States of America

Ryusuke Niwa

ryusuke-niwa@umin.ac.jp

Affiliations Graduate School of Life and Environmental Sciences, University of Tsukuba, Tsukuba, Ibaraki, Japan, PRESTO, Japan Science and Technology Agency, Kawaguchi, Saitama, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: TKK KH YSN MBO RN. Performed the experiments: TKK KH YSN RY MS RN. Analyzed the data: TKK KH YSN RY MBO RN. Contributed reagents/materials/analysis tools: TKK KH YSN MS TS AF MBO RN. Wrote the paper: TKK KH YSN MBO RN.