About the Authors

Keiko Myouzen

Affiliation Laboratory for Autoimmune Diseases, Center for Genomic Medicine (CGM), RIKEN, Yokohama, Japan

Yuta Kochi

ykochi@src.riken.jp

Affiliations Laboratory for Autoimmune Diseases, Center for Genomic Medicine (CGM), RIKEN, Yokohama, Japan, Department of Allergy and Rheumatology, Graduate School of Medicine, the University of Tokyo, Tokyo, Japan

Yukinori Okada

Affiliations Laboratory for Autoimmune Diseases, Center for Genomic Medicine (CGM), RIKEN, Yokohama, Japan, Department of Allergy and Rheumatology, Graduate School of Medicine, the University of Tokyo, Tokyo, Japan, Laboratory for Statistical Analysis, CGM, RIKEN, Yokohama, Japan

Chikashi Terao

Affiliations Center for Genomic Medicine, Kyoto University Graduate School of Medicine, Kyoto, Japan, Department of Rheumatology and Clinical Immunology, Graduate School of Medicine, Kyoto University, Kyoto, Japan

Akari Suzuki

Affiliation Laboratory for Autoimmune Diseases, Center for Genomic Medicine (CGM), RIKEN, Yokohama, Japan

Katsunori Ikari

Affiliation Institute of Rheumatology, Tokyo Women's Medical University, Tokyo, Japan

Tatsuhiko Tsunoda

Affiliation Laboratory for Medical Informatics, CGM, RIKEN, Yokohama, Japan

Atsushi Takahashi

Affiliation Laboratory for Statistical Analysis, CGM, RIKEN, Yokohama, Japan

Michiaki Kubo

Affiliation Laboratory for Genotyping Development, CGM, RIKEN, Yokohama, Japan

Atsuo Taniguchi

Affiliation Institute of Rheumatology, Tokyo Women's Medical University, Tokyo, Japan

Fumihiko Matsuda

Affiliations Center for Genomic Medicine, Kyoto University Graduate School of Medicine, Kyoto, Japan, CREST Program, Japan Science and Technology Agency, Kawaguchi, Saitama, Japan, Institut National de la Sant? et de la Recherche M?dicale (INSERM), Unit? U852, Kyoto University Graduate School of Medicine, Kyoto, Japan

Koichiro Ohmura

Affiliation Department of Rheumatology and Clinical Immunology, Graduate School of Medicine, Kyoto University, Kyoto, Japan

Shigeki Momohara

Affiliation Institute of Rheumatology, Tokyo Women's Medical University, Tokyo, Japan

Tsuneyo Mimori

Affiliation Department of Rheumatology and Clinical Immunology, Graduate School of Medicine, Kyoto University, Kyoto, Japan

Hisashi Yamanaka

Affiliation Institute of Rheumatology, Tokyo Women's Medical University, Tokyo, Japan

Naoyuki Kamatani

Affiliation Laboratory for International Alliance, CGM, RIKEN, Yokohama, Japan

Ryo Yamada

Affiliation Unit of Statistical Genetics, Center for Genomic Medicine, Graduate School of Medicine, Kyoto University, Kyoto, Japan

Yusuke Nakamura

Affiliation Laboratory of Molecular Medicine, Human Genome Center, Institute of Medical Science, University of Tokyo, Tokyo, Japan

Kazuhiko Yamamoto

Affiliations Laboratory for Autoimmune Diseases, Center for Genomic Medicine (CGM), RIKEN, Yokohama, Japan, Department of Allergy and Rheumatology, Graduate School of Medicine, the University of Tokyo, Tokyo, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: K Myouzen, Y Kochi, Y Okada, C Terao, K Ikari, K Ohmura, R Yamada, K Yamamoto. Performed the experiments: K Myouzen, Y Kochi, C Terao, A Suzuki, K Ikari, K Ohmura. Analyzed the data: K Myouzen, Y Kochi, Y Okada, C Terao, T Tsunoda, A Takahashi, R Yamada. Contributed reagents/materials/analysis tools: M Kubo, A Taniguchi, F Matsuda, K Ohmura, S Momohara, T Mimori, H Yamanaka, N Kamatani, Y Nakamura. Wrote the paper: K Myouzen, Y Kochi, Y Okada, C Terao, K Yamamoto.