About the Authors

Alex Yick-Lun So

Affiliations Department of Cellular and Molecular Pharmacology, University of California San Francisco, San Francisco, California, United States of America , Chemistry and Chemical Biology Graduate Program, University of California San Francisco, San Francisco, California, United States of America

Christina Chaivorapol

Affiliations Department of Biochemistry and Biophysics, University of California San Francisco, San Francisco, California, United States of America , California Institute for Quantitative Biomedical Research, University of California San Francisco, San Francisco, California, United States of America , Graduate Program in Biological and Medical Informatics, University of California San Francisco, San Francisco, California, United States of America

Eric C Bolton

Affiliation Department of Cellular and Molecular Pharmacology, University of California San Francisco, San Francisco, California, United States of America

Hao Li

Affiliations Department of Biochemistry and Biophysics, University of California San Francisco, San Francisco, California, United States of America , California Institute for Quantitative Biomedical Research, University of California San Francisco, San Francisco, California, United States of America , Graduate Program in Biological and Medical Informatics, University of California San Francisco, San Francisco, California, United States of America

Keith R Yamamoto

To whom correspondence should be addressed. E-mail: yamamoto@cmp.ucsf.edu

Affiliations Department of Cellular and Molecular Pharmacology, University of California San Francisco, San Francisco, California, United States of America , Chemistry and Chemical Biology Graduate Program, University of California San Francisco, San Francisco, California, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

AYLS, CC, ECB, HL, and KRY conceived and designed the experiments. AYLS, CC, and ECB performed the experiments. This manuscript was written by AYLS, CC, and KRY with guidance from ECB and HL.