About the Authors

Cécile Libioulle

Affiliation Unit of Animal Genomics, GIGA-R and Faculty of Veterinary Medicine, University of Liège, Liège, Belgium

Edouard Louis

Affiliation Unit of Hepatology and Gastroenterology, Department of Clinical Sciences, GIGA-R, Faculty of Medicine, and CHU de Liège, University of Liège, Liège, Belgium

Sarah Hansoul

Affiliation Unit of Animal Genomics, GIGA-R and Faculty of Veterinary Medicine, University of Liège, Liège, Belgium

Cynthia Sandor

Affiliation Unit of Animal Genomics, GIGA-R and Faculty of Veterinary Medicine, University of Liège, Liège, Belgium

Frédéric Farnir

Affiliation Unit of Animal Genomics, GIGA-R and Faculty of Veterinary Medicine, University of Liège, Liège, Belgium

Denis Franchimont

Affiliation Department of Gastroenterology, Erasmus Hospital, Free University of Brussels, Brussels, Belgium

Séverine Vermeire

Affiliation Department of Gastroenterology, University Hospital Leuven, Leuven, Belgium

Olivier Dewit

Affiliation Department of Gastroenterology, Clinique Universitaire St Luc, UCL, Brussels, Belgium

Martine de Vos

Affiliation Department of Hepatology and Gastroenterology, Ghent University Hospital, Ghent, Belgium

Anna Dixon

Affiliation National Heart and Lung Institute, Imperial College London, England

Bruno Demarche

Affiliation Unit of Bioinformatics, GIGA-R and Institut Montefiore, University of Liège, Liège, Belgium

Ivo Gut

Affiliation Centre National de Génotypage, Evry, France

Simon Heath

Affiliation Centre National de Génotypage, Evry, France

Mario Foglio

Affiliation Centre National de Génotypage, Evry, France

Liming Liang

Affiliation Centre for Statistical Genetics, Department of Biostatistics, Ann Arbor, Michigan, United States

Debby Laukens

Affiliation Department of Hepatology and Gastroenterology, Ghent University Hospital, Ghent, Belgium

Myriam Mni

Affiliation Unit of Animal Genomics, GIGA-R and Faculty of Veterinary Medicine, University of Liège, Liège, Belgium

Diana Zelenika

Affiliation Centre National de Génotypage, Evry, France

André Van Gossum

Affiliation Department of Gastroenterology, Erasmus Hospital, Free University of Brussels, Brussels, Belgium

Paul Rutgeerts

Affiliation Department of Gastroenterology, University Hospital Leuven, Leuven, Belgium

Jacques Belaiche

Affiliation Unit of Hepatology and Gastroenterology, Department of Clinical Sciences, GIGA-R, Faculty of Medicine, and CHU de Liège, University of Liège, Liège, Belgium

Mark Lathrop

Affiliation Centre National de Génotypage, Evry, France

Michel Georges

To whom correspondence should be addressed. E-mail: michel.georges@ulg.ac.be

Affiliation Unit of Animal Genomics, GIGA-R and Faculty of Veterinary Medicine, University of Liège, Liège, Belgium

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

E. Louis, M. Lathrop, and M. Georges conceived and designed the experiments. C. Libioulle, A. Dixon, I. Gut, M. Mni, and D. Zelenika performed the experiments. C. Libioulle, E. Louis, S. Hansoul, C. Sandor, F. Farnir, B. Demarche, M. Foglio, S. Heath, L. Liang, M. Lathrop, and M. Georges analyzed the data. E. Louis, D. Franchimont, S. Vermeire, O. Dewit, M. de Vos, D. Laukens, A. Van Gossum, P. Rutgeerts, and J. Belaiche contributed reagents/materials/analysis tools. M. Lathrop and M. Georges wrote the paper.