About the Authors

Yang Li

Contributed equally to this work with: Yang Li, Olga Alda Álvarez

Affiliation Groningen Bioinformatics Centre, Groningen Biomolecular Sciences and Biotechnology Institute, University of Groningen, Haren, The Netherlands

Olga Alda Álvarez

Contributed equally to this work with: Yang Li, Olga Alda Álvarez

Affiliation Laboratory of Nematology, Wageningen University, Wageningen, The Netherlands

Evert W Gutteling

Affiliation Laboratory of Nematology, Wageningen University, Wageningen, The Netherlands

Marcel Tijsterman

Affiliation Hubrecht Laboratory, Netherlands Institute of Developmental Biology, Utrecht, The Netherlands

Jingyuan Fu

Affiliation Groningen Bioinformatics Centre, Groningen Biomolecular Sciences and Biotechnology Institute, University of Groningen, Haren, The Netherlands

Joost A. G Riksen

Affiliation Laboratory of Nematology, Wageningen University, Wageningen, The Netherlands

Esther Hazendonk

Affiliation Hubrecht Laboratory, Netherlands Institute of Developmental Biology, Utrecht, The Netherlands

Pjotr Prins

Affiliations Groningen Bioinformatics Centre, Groningen Biomolecular Sciences and Biotechnology Institute, University of Groningen, Haren, The Netherlands , Laboratory of Nematology, Wageningen University, Wageningen, The Netherlands

Ronald H. A Plasterk

Affiliation Hubrecht Laboratory, Netherlands Institute of Developmental Biology, Utrecht, The Netherlands

Ritsert C Jansen

To whom correspondence should be addressed. E-mail: R.C.Jansen@rug.nl (RCJ), Jan.Kammenga@wur.nl (JEK)

Affiliation Groningen Bioinformatics Centre, Groningen Biomolecular Sciences and Biotechnology Institute, University of Groningen, Haren, The Netherlands

Rainer Breitling

Affiliation Groningen Bioinformatics Centre, Groningen Biomolecular Sciences and Biotechnology Institute, University of Groningen, Haren, The Netherlands

Jan E Kammenga

To whom correspondence should be addressed. E-mail: R.C.Jansen@rug.nl (RCJ), Jan.Kammenga@wur.nl (JEK)

Affiliation Laboratory of Nematology, Wageningen University, Wageningen, The Netherlands

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

YL analyzed and interpreted data and wrote the manuscript. OAÁ performed the microarray experiment. EWG created RIL and genotyping. MT performed SNP mapping and RIL generation. JF performed the distant pair design and microarray normalization. JAGR contributed to the RIL generation and genotyping and performed the microarray experiment. EH performed SNP mapping and genotyping. PP helped with the microarray analysis. RHAP supervised with SNP mapping and RIL design. RB contributed to the statistical analysis and wrote the manuscript. RCJ and JEK initiated and supervised the study and revised the manuscript.