About the Authors

David Garrick

Affiliation MRC Molecular Haematology Unit, Weatherall Institute of Molecular Medicine, University of Oxford, Oxford, United Kingdom

Jackie A Sharpe

Affiliation MRC Molecular Haematology Unit, Weatherall Institute of Molecular Medicine, University of Oxford, Oxford, United Kingdom

Ruth Arkell

Affiliation MRC Mammalian Genetics Unit, Harwell, Oxfordshire, United Kingdom

Lorraine Dobbie

Affiliation Institute for Stem Cell Research, University of Edinburgh, Edinburgh, United Kingdom

Andrew J. H Smith

Affiliation Institute for Stem Cell Research, University of Edinburgh, Edinburgh, United Kingdom

William G Wood

Affiliation MRC Molecular Haematology Unit, Weatherall Institute of Molecular Medicine, University of Oxford, Oxford, United Kingdom

Douglas R Higgs

Affiliation MRC Molecular Haematology Unit, Weatherall Institute of Molecular Medicine, University of Oxford, Oxford, United Kingdom

Richard J Gibbons

To whom correspondence should be addressed. E-mail: richard.gibbons@molecular-medicine.oxford.ac.uk

Affiliation MRC Molecular Haematology Unit, Weatherall Institute of Molecular Medicine, University of Oxford, Oxford, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

DG, AJHS, WGW, DRH, and RJG conceived and designed the experiments. DG, JAS, RA, and RJG performed the experiments. DG, RA, WGW, DRH, and RJG analysed the data. LD contributed reagents/materials/analysis tools. LD provided technical assistance. DG wrote the paper.