About the Authors

James Ronald

Affiliations Department of Genome Sciences, University of Washington, Seattle, Washington, United States of America , Fred Hutchinson Cancer Research Center, Seattle, Washington, United States of America

Rachel B Brem

Affiliation Fred Hutchinson Cancer Research Center, Seattle, Washington, United States of America

Jacqueline Whittle

address: Infectious Disease Research Institute, Seattle, Washington, United States of America

Affiliations Fred Hutchinson Cancer Research Center, Seattle, Washington, United States of America , Howard Hughes Medical Institute, Seattle, Washington, United States of America

Leonid Kruglyak

To whom correspondence should be addressed. E-mail: leonid@genomics.princeton.edu

Affiliations Fred Hutchinson Cancer Research Center, Seattle, Washington, United States of America , Howard Hughes Medical Institute, Seattle, Washington, United States of America , Lewis–Sigler Institute for Integrative Genomics and Department of Ecology and Evolutionary Biology, Princeton University, Princeton, New Jersey, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

JR, RBB, and LK conceived and designed the experiments. JR and JW performed the experiments. JR, RBB, and LK analyzed the data and wrote the paper.