About the Authors

Phillip Phuc Le

Affiliation Department of Genetics, University of Pennsylvania School of Medicine, Philadelphia, Pennsylvania, United States of America

Joshua R Friedman

Affiliations Department of Genetics, University of Pennsylvania School of Medicine, Philadelphia, Pennsylvania, United States of America , Department of Pediatrics, University of Pennsylvania School of Medicine, Philadelphia, Pennsylvania, United States of America

Jonathan Schug

Affiliation Center for Bioinformatics, University of Pennsylvania School of Medicine, Philadelphia, Pennsylvania, United States of America

John E Brestelli

Affiliation Department of Genetics, University of Pennsylvania School of Medicine, Philadelphia, Pennsylvania, United States of America

J. Brandon Parker

Affiliation Department of Genetics, University of Pennsylvania School of Medicine, Philadelphia, Pennsylvania, United States of America

Irina M Bochkis

Affiliation Department of Genetics, University of Pennsylvania School of Medicine, Philadelphia, Pennsylvania, United States of America

Klaus H Kaestner

To whom correspondence should be addressed. E-mail: Kaestner@mail.med.upenn.edu

Affiliation Department of Genetics, University of Pennsylvania School of Medicine, Philadelphia, Pennsylvania, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

PPL, JRF, and KHK conceived and designed the experiments. PPL, JRF, JEB, JBP, and IMB performed the experiments. PPL, JS, JEB, and IMB analyzed the data. JS contributed reagents/materials/analysis tools. PPL, JRF, and KHK wrote the paper.