About the Authors

Felix L. Wu

Roles Data curation, Formal analysis, Investigation, Validation, Visualization, Writing – original draft, Writing – review & editing

flw2113@cumc.columbia.edu (FLW); mp3284@columbia.edu (MP)

Affiliations Department of Systems Biology, Columbia University, New York, New York, United States of America, Integrated Program in Cellular, Molecular, and Biomedical Studies, Columbia University, New York, New York, United States of America

Alva I. Strand

Roles Data curation, Formal analysis, Investigation, Writing – review & editing

Current address: Department of Biology, Corix Plains Institute, Oklahoma Biological Survey, University of Oklahoma, Norman, Oklahoma, United States of America

Affiliation Department of Biological Sciences, Columbia University, New York, New York, United States of America

Laura A. Cox

Roles Funding acquisition, Resources, Writing – review & editing

Affiliations Center for Precision Medicine, Department of Internal Medicine, Section of Molecular Medicine, Wake Forest School of Medicine, Winston-Salem, North Carolina, United States of America, Southwest National Primate Research Center, Texas Biomedical Research Institute, San Antonio, Texas, United States of America

Carole Ober

Roles Data curation, Funding acquisition, Resources, Writing – review & editing

Affiliation Department of Human Genetics, The University of Chicago, Chicago, Illinois, United States of America

Jeffrey D. Wall

Roles Data curation, Funding acquisition, Resources, Writing – review & editing

Affiliation Institute for Human Genetics, Department of Epidemiology & Biostatistics, University of California, San Francisco, San Francisco, California, United States of America

Priya Moorjani

Contributed equally to this work with: Priya Moorjani, Molly Przeworski

Roles Formal analysis, Funding acquisition, Investigation, Methodology, Supervision, Writing – review & editing

Current address: Department of Molecular and Cell Biology, Center for Computational Biology, University of California, Berkeley, Berkeley, California, United States of America

Affiliation Department of Biological Sciences, Columbia University, New York, New York, United States of America

Molly Przeworski

Contributed equally to this work with: Priya Moorjani, Molly Przeworski

Roles Conceptualization, Funding acquisition, Investigation, Methodology, Project administration, Resources, Supervision, Writing – original draft, Writing – review & editing

flw2113@cumc.columbia.edu (FLW); mp3284@columbia.edu (MP)

Affiliations Department of Systems Biology, Columbia University, New York, New York, United States of America, Department of Biological Sciences, Columbia University, New York, New York, United States of America

Competing Interests

The authors have declared that no competing interests exist.