About the Authors

Natasha Saint Savage

Affiliation Department of Molecular Biology and Biotechnology, University of Sheffield, Sheffield, United Kingdom

Tom Walker

Affiliation Department of Cell and Developmental Biology, John Innes Centre, Norwich, United Kingdom

Yana Wieckowski

Affiliation Department of Molecular, Cell, and Developmental Biology, University of Michigan, Ann Arbor, Michigan, United States of America

John Schiefelbein

Affiliation Department of Molecular, Cell, and Developmental Biology, University of Michigan, Ann Arbor, Michigan, United States of America

Liam Dolan

Affiliation Department of Cell and Developmental Biology, John Innes Centre, Norwich, United Kingdom

Nicholas A. M Monk

To whom correspondence should be addressed. E-mail: nick.monk@nottingham.ac.uk

Affiliations Division of Applied Mathematics, School of Mathematical Sciences, University of Nottingham, Nottingham, United Kingdom , Centre for Plant Integrative Biology, School of Biosciences, University of Nottingham, Loughborough, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

NSS, TW, JS, LD, and NAMM conceived and designed the experiments. NSS, TW, YW, and NAMM performed the experiments. NSS and NAMM analyzed the data. NSS, LD, and NAMM wrote the paper.