About the Authors

Sejin Ahn

Contributed equally to this work with: Sejin Ahn, Gwanghee Lee, Soo Jung Yang, Deokjae Lee

Current address: Program in Biological and Biomedical Sciences, Harvard Medical School, Boston, Massachusetts, United States of America

Affiliation Laboratory of Cellular and Molecular Immunology, National Institute of Allergy and Infectious Diseases, Bethesda, Maryland, United States of America

Gwanghee Lee

Contributed equally to this work with: Sejin Ahn, Gwanghee Lee, Soo Jung Yang, Deokjae Lee

Current address: Department of Cell Biology and Physiology, Washington University School of Medicine, St. Louis, Missouri, United States of America

Affiliation Laboratory of Cellular and Molecular Immunology, National Institute of Allergy and Infectious Diseases, Bethesda, Maryland, United States of America

Soo Jung Yang

Contributed equally to this work with: Sejin Ahn, Gwanghee Lee, Soo Jung Yang, Deokjae Lee

Current address: Division of Biological Sciences, Seoul National University, Seoul, Korea

Affiliation Laboratory of Cellular and Molecular Immunology, National Institute of Allergy and Infectious Diseases, Bethesda, Maryland, United States of America

Deokjae Lee

Contributed equally to this work with: Sejin Ahn, Gwanghee Lee, Soo Jung Yang, Deokjae Lee

Affiliation Research Center for Molecular and Cellular Biology, Inha University, Incheon, Korea

Seunghyuk Lee

Affiliation Department of Biological Sciences, Inha University, Incheon, Korea

Hyo Sun Shin

Affiliation Department of Biological Sciences, Inha University, Incheon, Korea

Min Cheol Kim

Affiliation Department of Biological Sciences, Inha University, Incheon, Korea

Kee Nyung Lee

Current address: Laboratory of Immunology, Korea Research Institute of Bioscience and Biotechnology, Yusong, Taejon, Korea

Affiliation Laboratory of Cellular and Molecular Immunology, National Institute of Allergy and Infectious Diseases, Bethesda, Maryland, United States of America

Douglas C Palmer

Affiliation National Institute of Cancer, National Institutes of Health, Bethesda, Maryland, United States of America

Marc R Theoret

Affiliation National Institute of Cancer, National Institutes of Health, Bethesda, Maryland, United States of America

Eric J Jenkinson

Affiliation Medical Research Council (MRC) Centre for Immune Regulation, Institute for Biomedical Research, University of Birmingham, United Kingdom

Graham Anderson

Affiliation Medical Research Council (MRC) Centre for Immune Regulation, Institute for Biomedical Research, University of Birmingham, United Kingdom

Nicholas P Restifo

Affiliation National Institute of Cancer, National Institutes of Health, Bethesda, Maryland, United States of America

Moon Gyo Kim

To whom correspondence should be addressed. E-mail: mgkim@inha.ac.kr

Affiliations Research Center for Molecular and Cellular Biology, Inha University, Incheon, Korea , Department of Biological Sciences, Inha University, Incheon, Korea

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

MGK conceived and designed the experiments. SA, GL, SJY, DL, HSS, MCK, and MGK performed the experiments. SA, GL, SJY, DL, SL, HSS, MCK, EJJ, GA, and MGK analyzed the data. KNL, DCP, MRT, EJJ, GA, NPR, and MGK contributed reagents/materials/analysis tools. SA, GL, SJY, DL, SL, DCP, GA, NPR, and MGK wrote the paper.