About the Authors

Jason H Brickner

To whom correspondence should be addressed. E-mail: jbrick@itsa.ucsf.edu

Affiliation Howard Hughes Medical Institute and the Department of Biochemistry and Biophysics, University of California, San Francisco, California, United States of America

Peter Walter

Affiliation Howard Hughes Medical Institute and the Department of Biochemistry and Biophysics, University of California, San Francisco, California, United States of America

Competing Interests

The authors have declared that no conflicts of interest exist.

Author Contributions

JHB and PW conceived of the experiments. JHB performed the experiments. JHB and PW wrote the paper.