About the Authors

Atiyeh Peiravan

Roles Data curation, Formal analysis, Investigation, Project administration, Resources, Visualization, Writing – original draft

apeiravan@rvc.ac.uk

Affiliation Department of Pathology and Pathogen Biology, Royal Veterinary College, University of London, North Mymms, United Kingdom

Francesca Bertolini

Roles Formal analysis, Investigation, Visualization, Writing – review & editing

Affiliation Department of Animal Science, Iowa State University, Ames, Iowa, United States of America

Max F. Rothschild

Roles Formal analysis, Resources, Visualization, Writing – review & editing

Affiliation Department of Animal Science, Iowa State University, Ames, Iowa, United States of America

Kenneth W. Simpson

Roles Investigation

Affiliation College of Veterinary Medicine, Cornell University, Ithaca, NY, United States of America

Albert E. Jergens

Roles Investigation

Affiliation College of Veterinary Medicine, Iowa State University, Ames, Iowa, United States of America

Karin Allenspach

Roles Project administration, Resources, Supervision, Writing – review & editing

‡ These authors are shared senior authors on this work.

Affiliation College of Veterinary Medicine, Iowa State University, Ames, Iowa, United States of America

Dirk Werling

Roles Investigation, Project administration, Resources, Supervision, Writing – review & editing

‡ These authors are shared senior authors on this work.

Affiliation Department of Pathology and Pathogen Biology, Royal Veterinary College, University of London, North Mymms, United Kingdom

Competing Interests

We have the following interests: This study was funded in part by Laboklin GmBH. There are no patents, products in development or marketed products to declare. This does not alter our adherence to all the PLOS ONE policies on sharing data and materials.