About the Authors

Jennifer B. Nagashima

Affiliations Baker Institute for Animal Health, College of Veterinary Medicine, Cornell University, Ithaca, New York, United States of America, Atkinson Center for a Sustainable Future, Cornell University, Ithaca, New York, United States of America, Smithsonian Conservation Biology Institute, National Zoological Park, Front Royal, Virginia, United States of America

Skylar R. Sylvester

Affiliation Baker Institute for Animal Health, College of Veterinary Medicine, Cornell University, Ithaca, New York, United States of America

Jacquelyn L. Nelson

Affiliation Baker Institute for Animal Health, College of Veterinary Medicine, Cornell University, Ithaca, New York, United States of America

Soon Hon Cheong

Affiliation Department of Clinical Sciences, College of Veterinary Medicine, Cornell University, Ithaca, New York, United States of America

Chinatsu Mukai

Affiliation Baker Institute for Animal Health, College of Veterinary Medicine, Cornell University, Ithaca, New York, United States of America

Colleen Lambo

Affiliation Baker Institute for Animal Health, College of Veterinary Medicine, Cornell University, Ithaca, New York, United States of America

James A. Flanders

Affiliation Department of Clinical Sciences, College of Veterinary Medicine, Cornell University, Ithaca, New York, United States of America

Vicki N. Meyers-Wallen

Affiliation Baker Institute for Animal Health, College of Veterinary Medicine, Cornell University, Ithaca, New York, United States of America

Nucharin Songsasen

Affiliation Smithsonian Conservation Biology Institute, National Zoological Park, Front Royal, Virginia, United States of America

Alexander J. Travis

ajt32@cornell.edu

Affiliations Baker Institute for Animal Health, College of Veterinary Medicine, Cornell University, Ithaca, New York, United States of America, Atkinson Center for a Sustainable Future, Cornell University, Ithaca, New York, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JBN SRS CM CL VNMW NS AJT. Performed the experiments: JBN SRS JLN SHC CM CL JAF. Analyzed the data: JBN SRS JLN CM AJT. Wrote the paper: JBN SRS JLN SHC CM CL JAF VNMW NS AJT. Taught and performed techniques: SHC CM CL JAF NS.