About the Authors

Douglas F. Bertram

Contributed equally to this work with: Douglas F. Bertram, Mark C. Drever, Murdoch K. McAllister

douglas.bertram@dfo-mpo.gc.ca

Affiliation Environment Canada, Wildlife Research Division, Institute of Ocean Sciences, Sidney, British Columbia, Canada

Mark C. Drever

Contributed equally to this work with: Douglas F. Bertram, Mark C. Drever, Murdoch K. McAllister

Affiliation Environment Canada, Canadian Wildlife Service, Delta, British Columbia, Canada

Murdoch K. McAllister

Contributed equally to this work with: Douglas F. Bertram, Mark C. Drever, Murdoch K. McAllister

Affiliation Fisheries Centre, University of British Columbia, Vancouver, British Columbia, Canada

Bernard K. Schroeder

Affiliation Bernard K. Schroeder Consulting, Nanaimo, British Columbia, Canada

David J. Lindsay

Affiliation TimberWest Forest Corp., Nanaimo, British Columbia, Canada

Deborah A. Faust

Affiliation Kingston, Ontario, Canada

Competing Interests

DL is an employee of TimberWest Forest Corp., whose company funded this study. BKS is owner of Bernard K. Schroeder Consulting. There are no patents, products in development or marketed products to declare. This does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: DFB DJL BKS. Performed the experiments: BKS DJL. Analyzed the data: MCD MKM DFB DAF. Contributed reagents/materials/analysis tools: MKM MCD DAF. Wrote the paper: DFB MCD MKM BKS DJL DAF.