About the Authors

Yayoi Yoshimura

Contributed equally to this work with: Yayoi Yoshimura, Akihiko Shiino

Affiliations Research Institute, Shiga Medical Center, Moriyama 5-4-30, Shiga 524–8524, Japan, Department of Neurosurgery, Shiga University of Medical Science, Shiga 520–2192, Japan

Akihiko Shiino

Contributed equally to this work with: Yayoi Yoshimura, Akihiko Shiino

Affiliations Biomedical MR Science Center, Shiga University of Medical Science, Shiga 520–2192, Japan, Department of Neurosurgery, Shiga University of Medical Science, Shiga 520–2192, Japan

Kazue Muraki

Affiliation Research Institute, Shiga Medical Center, Moriyama 5-4-30, Shiga 524–8524, Japan

Tadateru Fukami

Affiliation Department of Neurosurgery, Shiga University of Medical Science, Shiga 520–2192, Japan

Shigeki Yamada

Affiliation Department of Neurosurgery, Shiga Medical Center, Shiga 524–8524, Japan

Takeshi Satow

Affiliation Department of Neurosurgery, Shiga Medical Center, Shiga 524–8524, Japan

Miyuki Fukuda

Affiliation Department of Neurosurgery, Shiga Medical Center, Shiga 524–8524, Japan

Masaaki Saiki

Affiliation Department of Neurosurgery, Shiga Medical Center, Shiga 524–8524, Japan

Masato Hojo

Affiliation Department of Neurosurgery, Shiga Medical Center, Shiga 524–8524, Japan

Susumu Miyamoto

Affiliation Department of Neurosurgery, Graduate School of Medicine, Kyoto University, Kyoto 606–8507, Japan

Nobuyuki Onishi

Affiliation Division of Gene Regulation, School of Medicine, Keio University, Tokyo 160–8582, Japan

Hideyuki Saya

Affiliation Division of Gene Regulation, School of Medicine, Keio University, Tokyo 160–8582, Japan

Toshiro Inubushi

Affiliation Biomedical MR Science Center, Shiga University of Medical Science, Shiga 520–2192, Japan

Kazuhiko Nozaki

noz@belle.shiga-med.ac.jp (KN); tanigaki@res.med.shiga-pref.jp (KT)

Affiliation Department of Neurosurgery, Shiga University of Medical Science, Shiga 520–2192, Japan

Kenji Tanigaki

noz@belle.shiga-med.ac.jp (KN); tanigaki@res.med.shiga-pref.jp (KT)

Affiliation Research Institute, Shiga Medical Center, Moriyama 5-4-30, Shiga 524–8524, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: AS KN KT. Performed the experiments: YY AS KM TI KT. Analyzed the data: YY AS KM KT. Contributed reagents/materials/analysis tools: TF SY TS MF MS MH SM NO HS. Wrote the paper: YY KT.