About the Authors

Hideyuki Doi

doih@hiroshima-u.ac.jp

Affiliations LAFWEDY, Faculty of Agriculture, Ehime University, Matsuyama, Japan, Institute for Sustainable Sciences and Development, Hiroshima University, Higashi-Hiroshima, Japan

Kwang-Hyeon Chang

Affiliations Center for Marine Environmental Studies, Ehime University, Matsuyama, Japan, Department of Environmental Science and Engineering, Kyung Hee University, Seocheon-dong, Giheung-gu, Yongin-si, Korea

Yuichiro Nishibe

Affiliations Center for Marine Environmental Studies, Ehime University, Matsuyama, Japan, Atmosphere and Ocean Research Institute, The University of Tokyo, Kashiwa, Japan

Hiroyuki Imai

Affiliation LAFWEDY, Faculty of Agriculture, Ehime University, Matsuyama, Japan

Shin-ichi Nakano

Affiliations LAFWEDY, Faculty of Agriculture, Ehime University, Matsuyama, Japan, Center for Ecological Research, Kyoto University, Otsu, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: HD K-HC SN. Performed the experiments: HD K-HC YN HI SN. Analyzed the data: HD. Wrote the paper: HD K-HC YN SN.