About the Authors

Hirokazu Doi

Affiliation Department of Neurobiology and Behavior, Graduate School of Biomedical Sciences, Nagasaki University, Nagasaki, Japan

Shota Nishitani

Affiliation Department of Neurobiology and Behavior, Graduate School of Biomedical Sciences, Nagasaki University, Nagasaki, Japan

Takashi X. Fujisawa

Affiliation Department of Neurobiology and Behavior, Graduate School of Biomedical Sciences, Nagasaki University, Nagasaki, Japan

Tomoko Nagai

Affiliation Department of Neurobiology and Behavior, Graduate School of Biomedical Sciences, Nagasaki University, Nagasaki, Japan

Masaki Kakeyama

Affiliation Center for Disease Biology and Integrative Medicine, Graduate School and Faculty of Medicine, The University of Tokyo, Tokyo, Japan

Takahiro Maeda

Affiliation Department of Island and Community Medicine, Island Medical Research Institute, Graduate School of Biomedical Sciences, Nagasaki University, Nagasaki, Japan

Kazuyuki Shinohara

kazuyuki@nagasaki-u.ac.jp

Affiliation Department of Neurobiology and Behavior, Graduate School of Biomedical Sciences, Nagasaki University, Nagasaki, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: HD MK TM KS. Performed the experiments: HD SN TXF TN TM. Analyzed the data: HD SN MK. Contributed reagents/materials/analysis tools: HD MK. Wrote the paper: HD KS.