About the Authors

Daniel E. Impoinvil

d.impoinvil@liverpool.ac.uk

Affiliations LUCINDA Group, Department of Epidemiology and Population Health, Institute of Infection and Global Health, University of Liverpool, Leahurst Campus, Neston, Cheshire, United Kingdom, Brain Infections Group, Department of Clinical Infection, Microbiology and Immunology, Institute of Infection and Global Health, University of Liverpool, Liverpool, Merseyside, United Kingdom

Tom Solomon

Affiliation Brain Infections Group, Department of Clinical Infection, Microbiology and Immunology, Institute of Infection and Global Health, University of Liverpool, Liverpool, Merseyside, United Kingdom

W. William Schluter

Affiliation Programme for Immunization Preventable Diseases, World Health Organization Country Office for Nepal, Kathmandu, Nepal

Ajit Rayamajhi

Affiliations Brain Infections Group, Department of Clinical Infection, Microbiology and Immunology, Institute of Infection and Global Health, University of Liverpool, Liverpool, Merseyside, United Kingdom, Department of Pediatrics, National Academy of Medical Sciences, Kanti Children's Hospital, Maharajgunj, Kathmandu, Nepal

Ram Padarath Bichha

Affiliation Child Health Division, Department of Health Services, Ministry of Health and Population, Government of Nepal, Kathmandu, Nepal

Geeta Shakya

Affiliation National Public Health Laboratory, Department of Health Services, Ministry of Health and Population, Government of Nepal, Kathmandu, Nepal

Cyril Caminade

Affiliation Department of Earth's Changing Environment, School of Environmental Science, University of Liverpool, Liverpool, Merseyside, United Kingdom

Matthew Baylis

Affiliation LUCINDA Group, Department of Epidemiology and Population Health, Institute of Infection and Global Health, University of Liverpool, Leahurst Campus, Neston, Cheshire, United Kingdom

Competing Interests

Prof. Matthew Baylis is an Academic Editor for PLoS ONE.

Author Contributions

Conceived and designed the experiments: DEI CC MB. Performed the experiments: DEI. Analyzed the data: DEI. Wrote the paper: DEI MB TS WWS CC AR RPB GS. Conducted the laboratory testing and compilation of the data: GS RPB AR. Interpreted the results: DEI MB TS WWS.