About the Authors

David T. S. Hayman

dtsh2@cam.ac.uk

Affiliations Cambridge Infectious Diseases Consortium, University of Cambridge, Cambridge, United Kingdom, Institute of Zoology, Zoological Society of London, London, United Kingdom, Rabies and Wildlife Zoonoses Group, Veterinary Laboratories Agency, Weybridge, United Kingdom

Petra Emmerich

Affiliation Bernhard-Nocht Institute, Hamburg, Germany

Meng Yu

Affiliation CSIRO Livestock Industries, Australian Animal Health Laboratory, Geelong, Australia

Lin-Fa Wang

Affiliation CSIRO Livestock Industries, Australian Animal Health Laboratory, Geelong, Australia

Richard Suu-Ire

Affiliation Wildlife Division of the Ghana Forestry Commission, Accra, Ghana

Anthony R. Fooks

Affiliations Rabies and Wildlife Zoonoses Group, Veterinary Laboratories Agency, Weybridge, United Kingdom, National Centre for Zoonoses Research, University of Liverpool, Liverpool, United Kingdom

Andrew A. Cunningham

Affiliation Institute of Zoology, Zoological Society of London, London, United Kingdom

James L. N. Wood

Affiliation Cambridge Infectious Diseases Consortium, University of Cambridge, Cambridge, United Kingdom

Competing Interests

Prof Anthony R. Fooks is an Academic Editor for PLoS ONE.

Author Contributions

Conceived and designed the experiments: DTSH RSI ARF AAC JLNW. Performed the experiments: DTSH PE MY. Analyzed the data: DTSH LFW ARF. Contributed reagents/materials/analysis tools: PE ARF. Wrote the paper: DTSH LFW AAC JLNW.